The insect's head is sometimes referred to as the head-capsule, and is the insect's feeding and sensory centre. It supports the eyes, antennae and and jaws of the insect. (Note -: insects do not breath through their mouths but through their thoracic and abdominal spiracles) The upper-mid portion of an insects face is called the 'frons' below this is the 'clypeus' and below this the 'labrum' to either side of which may be seen the edges of the 'mandibles' in some insects various aspects of the 'maxilliary' palps may extend beyond and or below these even when viewed from front on.
The 'frons' = that area of the face below the top two 'ocelli' and above the 'frontoclypeal sulcus' (if and when this is visible) and in between the two 'frontogenal sulci', it supports the 'pharyngeal dilator' muscles and in immature forms it bears the lower two arms of the ecdysial cleavage lines. The 'clypeus' = that area of the face immediately below the frons (with which it may be fused in the absence of the frontoclypeal sulcus) and the frontoclypeal sulcus. It supports the 'cibarial dilator' muscles and may be divided horizontally into a 'post.' and 'anteclypeus'. The 'labrum' = is equivalent to the insect's upper lip and is generally moveable, it articulates with the clypeus by means of the 'clypeolabral suture'. The rest of the front of the head: that bit which is above the frons is known as the 'vertex'; the sides of the head are known as the 'gena'. For a more detailed, lateral (side) view of the insect head follow this link. The Insect Head: Lateral
![]() The Antennae
The antennae are an insects primary, non-visual, sense organs, though in a few rare instances they have become adapted for other purposes such as seizing prey items ( i.e. the larva of Chaoborus sp. {Diptera}) or holding females during mating (i.e. the males of Meloe sp. {Coleoptera})
The Eyes Though some species of insects have been shown to be able respond to light stimulus through their cuticle, most light sensitivity occurs through one or more eyes. Insects possess two different sorts of eyes, the usually large and obviously visible compound eyes, and two varieties of ocelli or simple eyes. Compound Eyes Compound eyes are so named because the cornea is composed of a number of individual facets or lenses (called ommatidia), rather than a single lens as in ocelli (or our own eyes). The number of separate visual elements or ommatidia varies greatly between species as well as between the larger taxa, so that while worker ants of different species may have between 1 (Ponera punctatissima) and 600 ommatidia per single eye, adult male Odonata may have more than 28,000 per single eye. Much like our eyes, the eyes of insects, can be divided into four basic parts: the supportative material that keeps all the parts together; a light gathering part (the lens and the auxilary lens called a 'crystalline cone'); a light receptor that converts the recieved light into electrical energy; and the nerves that carry the electrical impulses to the brain for analysis. In the compound eyes of insects these parts are repeated numerous times side by side in a space saving hexagonal pattern. See above. The lens is formed by a transparent and colourless cuticle and it is usually biconvex. Beneath this is the crystalline cone (which is comprised of four cells called 'Semper cells' after the man who first described them). Normally this functions as a secondary lens.
The corneal lens is supported by 'primary pigment cells' and the retinula cells and associated rhabdoms are supported by 'secondary pigment cells'. The retinula cells are connected to axons at the base of the eye, it is these which carry the information collected by the lenses and converted into electrical impulses by the rhabdom to the brain, thus allowing the insect to see. Simple Eyes
The function of the corneal lens is obscure, although it does project an image into the ocellus this image forms below the level of the light-sensitive cells, or rhabdom. Therefore the ocellus can generate no image information, however it is very sensitive to low levels of light and to changes in light intensity and scientists believe that the ocelli are useful in allowing the insect to detect the horizon, to respond quickly to changes in light intensity. Two different forms of ocelli have been described for insects, Dorsal ocelli and Lateral ocelli.
Smell
Most insects communicate using smell or chemoreception and it is not surprising that they have evolved a large variety of ways of detecting the molecules involved. Insects do not have noses like us which concentrate all our sense of smell in one place, instead they have a lot of small sensory bodies scattered over their body, though they tend to have a concentration of them on their antennae. We can recognise several different common forms of chemoreceptor though these are not the only forms they can take by any means,
The Mouthparts
In the primitive form the insect mouth articulates (moves) from side to side in a horizontal plane, rather than vertically as do ours. In those groups of insects that evolved later the basic mouthparts shown below have become highly modified, in the butterflies and moths they are transformed into a long flexible straw, in the blood and sap sucking insects of various orders they have become a hypodermic needle and in most of the flies you see from day to day they evolved into a extensible and highly effective sponge. The Mandibles
The Maxillae
The Labium results from the fusion of a pair of limbs and serves a purpose similar to our lower lip for the insects. The main body of the labium is divided into three parts the central 'mentum' which is boarded on either side by the 'submentum' proximally which hinges with the head, and the prementum distally. The prementum supports two pairs of lobes known as the 'glossae' and to the outside of them the 'paraglossae', and a pair of labial palps which are primarily sensory in function. The glossae and paraglossae may be fused, with one or the other considerably reduced, in which case the whole thing is known as the 'ligula'. The hyper-pharynx or tongue is found behind the mouth, and has the salivary ducts at its base, in most Diptera (true flies) and Hemiptera (true bugs) it has become highly modified and serves as the main feeding organ, in many cases combining with the rest of the mouthparts to form a stylette or piercing organ.
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